noc18-bt26-Lecture 05-Preliminaries

IIT Kanpur July 201
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Dec 3, 2023
Preliminaries
[Music]
[Music]
so let us now begin with our first
lecture of the module which is
preliminaries the first question that we
need to ask is why do we need to assess
the number of wildlife and there are a
number of reasons for that one numbers
are essential at every stage of
management now classically management is
said to follow the Deming cycle now
Deming cycle consists of four stages it
goes by the name of PD CA cycle which
means P is for plan followed by du
followed by check followed by Act and
followed by plan again now what do we
mean here plan means to take all the
steps and the decisions beforehand on
what we want to do why we want to do
that
and how are we going to do so when we
answer all these three questions this is
the planning stage the doing stage is
the actual execution of work now to take
an example suppose you have a forested
area which also has a thick undergrowth
of weed so one of our management
interventions could be to remove these
weeds so our aim is to remove weeds now
why do we want to remove these weeds
because these weeds are of a non
palatable species and because the
animals are not able to utilize this is
peace now so what we want to do is to
remove the weeds why we want to do it so
that more animals can utilize this space
and how are we going to do it maybe say
using a manual labor to uproot all these
weeds now once we have decided that once
we have decided on how much amount of
work is required at what rate we are
going to pay these laborers and things
like that will actually execute the work
on the field now while doing this
execution there might be some other
impacts that we had not thought of
beforehand so for instance when we went
there with our laborers and we saw that
most of the animals are just running
away from the field now this is
something that we had not expected
beforehand so which is why we'll come to
the checking stage so while we are doing
our work will be checking our work again
and again and again just to see whether
we are able to to reach our outcome of
the animals being able to utilize this
space in a more effective manner now
checking means that we are will look at
how many animals are utilizing this area
when we are doing this work and also
after we are done with our work and then
this check will be followed by act
now act actually means that we'll
utilize our knowledge of the checking
stage to incorporate it into our next
planning so for instance if you have a
last chunk of forest maybe we are doing
our weed removal in this a small patch
and once we have we are done with the
bead removal in this patch will utilize
the knowledge that we gained from our
exercise to plan the vida movin in
summary some other area so which is the
acting stage now numbers are crucial
inputted all of these stages because
when we said during our planning stage
that we wanted to remove these weeds so
that animals would be able to utilize
this area in a more effective manner and
because animals are not utilizing this
area in an effective manner because of
the presence of the weeds so in that
case what we need as our baseline
information is how many animals are
there in this weed infested area say in
a in a per hectare format while we are
doing our work while we are executing
our work will also incorporate data
about the weeds that are being removed
the rate at which we are removing the
weeds the time it is going to take to
remove these beads in a patch of area
and we'll also need information about
what the animals are doing in this
period of time in the checking stage so
once we are done with our weed removal
we again required data was this patch
once we have removed the weeds from
there is this patch being utilized by
the animals in a more effective manner
so essentially if we said if we found
out that our baseline with the presence
of feeds was say 15 animals per hectare
and after removal
after we have removed all these weeds
and after the animals have gotten inside
and are utilized in this area this
density moved up from 15 animals per
hectare to say 100 animals per hectare
now this is an information which will be
extremely crucial when we are doing the
next stage of planning
so essentially wildlife numbers are
required at every stage of Management
now second point why we need to assess
wildlife numbers is that numbers are
crucial inputs for decision support now
what does this mean
suppose we are managing a tiger reserve
and suppose we have the density of the
herbivores at say 50 animals per square
kilometers now is this 50 animals per
square kilometer good for our tigers in
the tiger reserve do we need more number
of animals do we need less number of
animals because if we have less number
of herbivores - less prey population so
our Tigers will not be able to reach
their optimal density so we will need to
increase the numbers of the herbivores
on the other hand if we have a situation
in which we have quite a lot of
herbivores in our area so we already
have a huge density of tigers which is
leading to more and more fights between
these tiles so we will need to reduce
our prey density now do we need to
increase our prey density or do we need
to reduce our prey density or do we need
to keep our prey density as constant
will able to gather information about it
only when we know their numbers so we
need to know the numbers of the
herbivores we need to know the numbers
of the tigers at the same time numbers
also help us assess the risk of a
population decline or a crash so as we
have seen beforehand in another lecture
if we have a population that is very
small will have a number of stochastic
phenomena that will become dominant in
that area now stochastic phenomena means
chance events so a chance event
essentially means that earlier if we had
say 1,000 animals
so suppose we had a huge population size
of our animals
suppose we had 10,000 animals and every
year this pool of 10,000 animals is
giving rise to say 1,000 young ones now
considering a sex ratio of one male is
to one female which is naturally found
in most of the situations we can say
that out of these 1,000 individuals see
around 500 would be males and we'll be
having 500 that are females and in in
such a scenario all of these animals
would be able to to mate properly
because they have a nice sex ratio but
considered we have only say 10 animals
left in our reserve and these 10 animals
gave rise to say two young ones and
these 10 animals are now growing old and
so our population growth is only
dependent on these two young ones now
even though we say that our sex ratio of
1 is to 1 would mean that we would have
on an average one male and one female
but because here we have such a small
population size it is also possible that
we got two males or two females now if
we have say two males and we do not have
any more females then this population
might be leading into a crash similarly
if we have only females left if you do
not have any means the population will
not be able to grow so these kinds of
phenomena are known as stochastic
phenomena
and at small population sizes we have
different stochastic phenomena as you
can see on your sites now so we have
demographic stochasticity now the
demographic stochasticity means demo is
your people graffia is to write
demographic stochasticity is a
stochastic city at the level of your
population numbers so here you can have
chance variations in births and deaths
leading to more deaths and less births
for instance so in a very small
population if more number of individuals
died out so you will be having a
population crash trans variations in sex
ratios as we saw in a population with
all males or all females or even
environmental stochasticity such as
draught flood famine or diseases now if
we have a large population of animals
then these stochastic these
environmental stochastic phenomena like
draught so floods do not have a major
impact because if we have say 10,000
animals and the draught killed as many
as say 5,000 animals we still have 5,000
animals left
but if we had a very small population
say of ten animals and if draught
occurred and say nine of these animals
died we'll be left only with one animal
which will again not be able to breed
and the population would be towards a
crash another stochastic phenomenon as
you can see on your sides now is genetic
problems such as inbreeding depression
now inbreeding depression occurs when
your animals are meeting with their
closest relatives
so for instance if you have a very small
population say you have only two
individuals left now these two
individuals after they have mated they
would produce an offspring and because
we have very less number of individuals
left now the next level of mating would
be say between the parents and and their
children or maybe between those children
themselves which would be a case of
inbreeding which also leads to a number
of genetic problems also genetic
problems could include genetic drift of
small populations so this is something
that will can we'll discuss in more
details in the conservation genetics
module and also the loss of
heterozygosity at the same time when you
have very small populations you also
have behavioral problems and ill effects
so for instance consider the case of
pack-hunting animals consider the case
of wild dogs now in the case of wild
dogs you have a large sized pack and say
ten to fifteen animals go to hunt at the
same time now in such a scenario if you
have a population in which you have only
five animals left and these animals are
adapted to pack hunting so they required
say 15 animals to hunt and you are left
only with five animals so these five
animals will not be able to hunt
sufficiently or efficiently enough so
that they would not be able to to gain
enough amount of food so this is another
stochastic phenomenon that will happen
another thing that happens when you have
a very small population size is that
the animals are highly dispersed they
are unable to find a mate so you have a
large size forest and in this forest you
have an a male here and you have a
female here but just because their
densities are so less they are not able
to find each other and so they are not
able to mate which generally have
happens when your population densities
are very less another situation when we
are required to assess wildlife numbers
is because numbers help us plan
scenarios and take steps so if we know
that our population is going down we can
take steps and these steps could include
adaptation steps and mitigation steps
now the difference between adaptation
and mitigation is that mitigation tries
to remove the root cause of the problem
whereas adaptation tries to acclimatize
your population so that even that even
when that root causes present you are
able to cope with it to give an example
from a very different field when we talk
about climate change adaptation and
mitigation so climate change mitigation
would mean that we'd reduce the amount
of carbon dioxide that we are putting
into that maas field or we increase the
amount of carbon dioxide that we are
taking out of the atmosphere so that is
mitigation because the root cause that
is the presence of carbon dioxide in the
air that is being addressed on the other
hand adaptation would mean that even
though we have carbon dioxide in the
atmosphere even though we are having
global warming we want our populations
to adapt to it so that would include
installation of say both number of air
conditioners is construction of more
number of dams so as to be able to cope
with the situations of draughts and
things like that so when we have numbers
we can plan scenarios even for our by
life and we can take steps for them
another question that we need to answer
is do we need numbers or do we need
trends so what do we mean by numbers and
trends so for instance in
our Tiger Reserve suppose we have 50
Tigers so this number 50 would be and
the number of Tigers but if we say that
say 10 years back we were having 70
Tigers and now we are having 50 Tigers
so we are having a downward trend so if
we do the number of Tigers versus the
years earlier we had 70 Tigers now we
have 50 Tigers
so this downward sloping graph would
give us the trend that this is a
decreasing trend on the other hand we
could also have an increasing trend so
earlier we had 70 Tigers now we have 90
Tigers at the same time we can also have
a trend of status quo in which earlier
also we had 70 Tigers now also we have
70 Tigers so do we need numbers or do we
need trends so actually we need both of
these why because in most of the
situation's it is easier to find out
trends and it is more difficult to find
out numbers so trains help us gather
information in a very short period of
time but numbers are also important
because they give us fine resolution of
data so for instance in our Tiger
Reserve we have tigers and we also have
herbivores in the case of herbivores it
does not matter much whether we have say
10,000 herbivores or we have 10,000 100
herbivores because the purpose of having
a large population of herbivores in our
Tiger Reserve is to act as a pre
population so in the case of herbivores
it would be worthwhile just to know
their trends whether this figure is
close to ten thousand or whether this
figure is going down drastically or
whether this figure is increasing
drastically so that course level of
information is good for us but for other
animals such as Tigers which are already
in a very small numbers we need to know
the actual number of tigers that are
there in that area
similarly with other animals such as the
Dewgong or the sea cow we or we have
only a few scores of sea cows left in
our ocean
and so it is extremely crucial that we
know the correct numbers and even a
single individual counts in that
scenario
another point to keep in mind between
numbers and trends is that with the
numbers you can always figure out the
trends but not vice versa
so for instance if I say that my trend
has been increasing for the last 20
years and now I have say 20 thousand
animals it is very difficult to to to
use just this data of an increasing
trend without knowing the actual numbers
to back calculate the number of animals
that were there beforehand but on the
other hand if we knew that the number of
animals in different years was say five
thousand five thousand five hundred six
thousand and say year 2000 2005 and 2010
so if we have these numbers where you
can very easily see that we are having
an increasing trend so it is very easy
to find out the trends from the numbers
and not vice-versa
now the next question is what are the
kinds of demographic information that we
are trying to get from our counting of
animals just a mere count of the number
of animals is not enough we also need to
know a number of other demographic
parameters one is the population size or
the number of individuals that are there
in the population and this is one
figured that we directly get out of our
counts from the population size if you
divide the size by the area you will get
the population density so it is
generally in per unit per hectare or per
square kilometer or in some animals
which have an even lesser density we we
generally make use of per hundred square
kilometers next is the H pyramid so the
H pyramid tells us the distribution of
various age groups in the population
which also tells us whether our
population is young population whether
it's an old population
whether it consists of a number of
adults because if our population
consists of mostly old animal spin it
might suffer a crash in the near future
then we have the crude birth in death
rates which tells us the number of life
or death or dead birth the the number of
live births or the number of deaths per
1000 individuals of the population we
also need to know the general fertility
rate so general fertility rate tells us
the the annual number of live births per
1000 females of reproductive age in the
population then we also need to know the
age specific fertility rate which is the
the annual number of live births per
1000 females of a specific age class in
the reproductive age so in this case we
can say for instance what is the age
specific fertility rate for females of
ages between 5 and 7 years in a tiger
population then next we have the total
fertility rate which is the average
number of live births per female
individual completing her reproductive
age if she followed the current age
specific fertility rate of the
population so essentially we are
integrating the age specific fertility
rate data to get the total fertility
rate the next very important information
is the replacement level fertility now
replacement level fertility means that
when you have a male and a female that
have mated how many offsprings are do
they need to produce so that they are
adequately replaced by the next
generation so the average number of
offsprings that a female individual must
produce such that the population is
completely replaced for the next
generation and the replacement level of
fertility is generally greater than or
equal to 2 now what this means is if you
have a male and a female and throughout
their lifetime they are only producing
one offspring so in that case if our
population in this generation as 1000
individuals in the next generation it
will become 500 individuals because out
of every 2 individuals there there is
only one individual that is being
produced on the other hand if you have a
replacement level of fertility which is
equal to two so that means that every
for every Bale and female they are
producing two offsprings so they are
getting completely replaced in the next
generation but then why do we say that
the replacement level of fertility is
greater than or equal to two now this is
because a number of offsprings
will die before reaching their
reproductive age so if you have juvenile
mortality or infant mortality so this
number would be greater than two now
juvenile mortality rate is the annual
number of deaths of juveniles per 1000
live births now here it is important to
note that this is one thousand live
births and not one thousand births life
expectancy is the number of years that
an average individual in the population
at a given age could expect to live at
the present age specific mortality
levels other information include
immigration which is how many
individuals are coming from outside into
our population emigration which is the
number of individuals that are going out
from our population into some other
population net migration which is
emigration - emigration naturally
increase which means in the case of
national increase we do not consider
migration into account
so naturally an increase is the number
of births minus the number of deaths
population growth which is births minus
deaths plus immigration minus emigration
so it's births plus immigration minus
deaths minus emigration so essentially
how does our population grow our
population will grow if we have any
births in the population or if there are
more animals that are coming from
outside
now our population would reduce if there
are deaths in the population or when the
animals are moving outside so when we
considered all of these four together we
get the population growth from
population growth we can also figure out
the population growth rate when we
divide it by the size of the population
over a fixed time period it is generally
expressed this percent per annum now the
next concept when we are estimating
animal numbers is absolute and relative
estimates of animal numbers now absolute
estimates of animal numbers
the size of the population or its
density by actually looking at the
animals or their cues such as nest so
examples include point counts and
discipline sampling and they generally
require more effort than relative
estimates and relative estimates on the
other hand do not count the actual
population rather they utilize proxies
to estimate animal densities in unknown
units so essentially what we want to say
in this case is that suppose you wanted
to figure out how many fishes are there
in a pond
now absolute estimate of the animal
number or the number of fishes would
actually involve taking a sample of
fishes from that area and counting the
number of pitches that are there in the
sample and then extrapolating it to the
whole of the area on the other hand a
relative estimate of the number of
fishes would include just say going into
that that pond fishing for say two words
and if I did a fishing now and suppose I
caught hundred fishes now I do this
experiment again I go to the pond again
and then I again fish for the same
amount of time and in pace of 100
individuals I gots a hundred in ten
individuals so a relative estimate of
animal numbers would say that the number
of animals in pond two is greater than
the number of animals in point one so
relative estimates permit comparisons
over space and time they're useful when
doing extensive works when actual number
of individuals in the population are not
required and rather trends are all that
we are looking after so let us now have
a look at some of the common techniques
of estimating population abundance so
some common techniques are complete
census now in a complete census you
count each and every individual of the
animal species that is present in your
area so this is generally utilized for
those animals that are already in very
small numbers so for instance in the
case of dugongs or the sea cows will
have a complete census of the number of
individuals that are there plot sampling
now in the case of a plot sampling what
we do
is that out of our complete forest we
take some small plots so they can be
rectangular plots or they could be
circular plots or they could be even be
our strip plots so we take some plots
suppose the areas are a1 a2 a3 and a4
and the total area of our region is
capital a now the number of animals that
we have in a1 as n1 here we have n2 here
we have n3 here we have n4 so we did not
count the total number of animals in our
complete forest but we only counted it
in four of the sample plots in that case
the total number of animals in the
sample plot which is n will be n1 plus
n2 plus n3 plus n4 the total area a of
all the four sample plots would be e 1
plus a 2 plus a 3 plus a 4 in which case
we'll get the density of animals which
is the number of animals per unit area
is given by sum of Ni divided by sum of
AI and then we'll extrapolate this
density to the whole area so we'll say
that the density of animals that we have
found using our 4 sample plots is the
same as the density of the animals and
our complete forest so we will have D is
equal to D now this small D we can
calculate because we have the n values
and we have the AI values and we have
this assumption that capital D which is
the density of animals and our complete
forest is the same as the density of
animals in the 4 sample plots in which
case to find out the total number of
animals in our forests we'll multiply
this capital T with the total area e to
get the number of animals so this is
plot sampling now distance sampling is
another technique in which which takes a
modification of a plot sampling and also
in
suits the data that not all the animals
are counted so there could be some
animals that we melt out during our
counting's and when we incorporate that
data in in our formula we get into the
distance sampling next we have the
capture mark recapture method sometimes
we cut curly Yampa so let us now
understand the capture mark recapture
technique by doing the small experiment
so in this jar we have some chickpeas
and we do not know the number of these
chickpeas so we'll perform the the
capture mark recapture method and I will
show you all the steps to to make an
estimate of the number of chickpeas that
are there in the jar so we first begin
by doing a capture of some animals so we
took out some animals these chickpeas
and here we have 1 2 3 4 5 10 15 20 so
we have 20 animals here let's take a few
more so we now have 25 animals so we
have captured these 25 animals next we
mark these animals so let us mark them
with a filter pin so I am marking them
with a black color
so in the field what we do is this
marking say in the case of birds would
be done by putting small bands into
their feet in the case of butterflies it
would be done by putting small stickers
on their wings in the case of ants it
would be done by spraying them with some
colored dyes and so on now the important
thing to note here is the number of
animals that we have captured if the
number of animals that we have captured
is very less then our errors in the
estimate would be great
whereas if the number of animals is more
then we'll have a much more correct
estimate of the total number of animals
in the forest so now that we have marked
all our 25 animals we now release them
back into the forest so we put them back
into the jar remember we had 25 animals
so 25 marked animals and then we let
them roam freely and make sure that the
mix with the other animals so now that
we have mixed these animals now let us
take out a sample so we do a recapture
of the enemies and now let us count the
number of animals that we have 5 6 7 8 9
10
15 20 25 30 35 40 45 47 50 54 now out of
these 54 animals we have 5 5 10 13
animals so we took out these samples of
54 animals and out of these 54 animals
we have 13 animals that are marked so
now let us do the calculations so let us
now look at the experiment again so in
the first case we took out n1 number of
animals which were 25 animals and we
marked these animals in the next stage
we took out into animals which is 54
animals and out of these 54 animals we
found that 13 animals were marked now if
we have 13 marked in 54 animals this
ratio would be same as 25 marked in the
total n animals that we have in our
forest because we are taking the
assumption that these animals have mixed
up completely they have not lost any
marks and we were able to see all these
marks very clearly so from this let us
find out what is n so n is equal to 54
into 25 divided by 13 so from here we
get the value of 103.8 animals so this
is the estimate
of number of animals in the forest which
in our case means the total number of
these chickpeas that were there in the
jar now I asked my associate to count
the number of chickpeas in the jar and
he came up with the figure of 100
chickpeas so there were actually 100
chickpeas and our estimate is one
hundred and three point eight which is a
very close estimate so coming back to
the slides captured mark-recapture
method is a very fast method of knowing
the total number of animals that are
there in our forest now another method
is the removal method now removal method
is not generally used in our country but
it is generally used in those countries
which perform hunting operations so
essentially if we take the data from
hunting and over the years the number of
hunts if it has remained constant so we
would say that our population has
remained constant but if with the same
amount of hunting effort our number of
hunts is going down then we would say
that the actual population is also going
down because of which we are getting a
lesser density of the animals so which
is briefly about the removal method but
it is not quite applicable to our
country because we do not permit hunting
so there are ways and means to decide
which technique to deploy in different
scenarios which is something that you
might have a look at but it is not
extremely crucial for this particular
course now the next concept is what is
the difference between precision and
accuracy so for instance if we are doing
any experiment if we repeat that
experiment again and again we'll be
getting different figures from that
experiment from those figures we can
figure out two things one is precision
one is accuracy and to understand that
let us look at the
results of this experiment so we have a
shooter and we have this bullseye and
these dots represent where these bullets
meant into the into the board now if we
look at this shooter so she fired five
shots all these five shots are centered
at the bullseye and all these five shots
are together now if we know the correct
value now the correct value in this case
should be the bullseye if the average of
our values is close to the correct value
we will see that the results are
accurate so here the results are
accurate here even though the shots are
very far off but still we can say that
the results are accurate because the
average of these five would fall
somewhere here but in this case the
results are not accurate because they
are at a quite a distance from the
actual correct figure which should be
this and in this case also the the
results are not accurate because we will
be seeing it at a distance so accuracy
is how close our result is to our
correct figure so basically in the case
of our capture mark-recapture experiment
we had the correct figure of hundred
animals but we got a figure of 103 point
eight animals from our experiment so our
figure is less than four percent away
from the actual correct figure so we
would say that our results are pretty
accurate now the next thing is about
precision precision is how close are our
values to each other so essentially if
you look at these values even though
these values are not accurate all these
five values are close together similarly
here also the five values are close
together so we will say that both of
these are precise results on the other
hand these values are very far off from
each other these values again are very
far off from each other so we would say
that both of these results are not
precise now coming back to our
experiment suppose we did this capture
mark-recapture
experiment again
and in place of 103.8 we got a figure of
say 102 or say 105 so we are doing these
experiments again and again and we are
getting these values of the first value
was 100 and 3.8 the second value was say
hundred and to 105 say 98 and 99 we did
this experiment for five times on the
other hand we did this experiment again
in another scenario the first figure 100
in 3.8 the second was say 150 then 250
and say 100 now in this case all these
values are close together the lowest
values 98 the highest value is 105 which
is a difference of only 7 points
in the second case our lowest value is
50 our highest value is 200 so the
difference here is 150 the difference
here is 7 so we would say that these
values are more precise as compared to
these values and coming to the accuracy
if we take the average of all these five
values if they are close to hundred we
would say that these values are accurate
else not so this is the difference
between precision and accuracy another
concept that we need to know is bias so
what is bias bias is the difference
between the mean of the measured values
and the reference value or the true
value so if the if the reference value
is the true value then the bias would
indicate the error of measurement so in
this case let us again go back to our
shooter
example so our shooter shot five shots
these shots are very precise because
they are all close together but they are
not accurate they have a difference be
between them and the correct value so
this difference B would be called the
bias now whenever we have a bias in our
results so in those situations what we
do is we calibrate
our instrument or we calibrate our
method so in this particular case the
shooter might be having a bend in her in
her gun so if and that bent in this case
is towards the right and towards the
upside so even though she was targeting
this point the results came here because
the gun had a bend like this so when we
calibrate our instrument we counter for
this bias and we ask our shooter to
shoot somewhere here so if she took a
target here she would come to the
correct value and similarly with all our
methods of measurement we would be
having some or the other bias which
would need to be calibrated so that's
all for today thank you for your
attention
[Music]
[Music]
